Managed to get a save where I could reproduce the issue.
https://drive.google.com/file/d/1bNJp_eP6BzKO4KyCKywbIoxCXbidaHT6/view?usp=sharing

Nests outside the South Eastern wall would respawn without regard to any settings/rules (no bugs present, no spawning in period, did not care about migration candidate chunks, did not matter if migration was disabled). Spawned in Nuclear Artillery/Ion Cannon to kill them quickly, and they would respawn within a minute as a group. 2 Screenshots about a minute apart are below.
Rampant 3.1.2 was installed at the start of the save.

https://imgur.com/a/b6ydrRK
Rampant 3.2.2
Evolution at .9594

Modlist:
https://drive.google.com/file/d/1UvBv2RkRCdlWwt3_ZxdChTD7plB5S1yi/view?usp=sharing

We've reverted to 3.1.2 for now without issues.

Came here to post this, the update has made it quite difficult to play Rampant. I suspected the new Compression feature to be the culprit but even after disabling it, I was having instantly respawning nests behind the lines.

Will confirm same issue, and same fix with rollback

This should be fixed in the latest