Seems like this mod makes Spitters not able to damage trees. Biters can damage trees fine when they run into a forest and get stuck, but Spitters will sit there indefinitely spitting at trees that it cannot destroy. Can confirm that turning this specific mod off allows them to damage the trees and move on.
Edit: Seems like it happens with rocks too. Clip of it happening here: https://streamable.com/1v9zqh